It is bordered in the north by Ecuador and Colombia , in the east by Brazil , in the southeast by Bolivia , in the south by Chile , and in the south and west by the Pacific Ocean. Peru is a megadiverse country with habitats ranging from the arid plains of the Pacific coastal region in the west to the peaks of the Andes mountains extending from the north to the southeast of the country to the tropical Amazon Basin rainforest in the east with the Amazon river. Peruvian territory was home to several ancient cultures. Ranging from the Norte Chico civilization starting in BCE, the oldest civilization in the Americas and one of the five cradles of civilization , to the Inca Empire , the largest state in the pre-Columbian Americas, the territory now including Peru has one of the longest histories of civilization of any country, tracing its heritage back to the 4th millennia BCE. The Spanish Empire conquered the region in the 16th century and established a viceroyalty that encompassed most of its South American territories, with its capital in Lima.
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